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GCSE level exam revision notes on Evolution: 8.

Evidence of human evolution - fossil skeletons, stone tools and modern genetic analysis

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[Key points and learning objectives for this page, after the main body of notes]

Sub-index of biology notes on all aspects of EVOLUTION


(8) The evidence of human evolution - from stone tools to modern genetic analysis

What does the fossil record and artefacts tell us about ourselves!

Since we are the species which has elucidated a theory of evolution, it seems appropriate to look at how we our selves (homo sapiens) have evolved!

As a child, I was fascinated by this picture of skeletons in a book my mother had, called "Apes and Men" published by Oxford: The Clarendon Press (Oxford University Press) in 1927.

The skeletons alone suggest they all have a common ancestor! but note the changes in the relative length of arms and legs, and the structure of the feet and skull (including the size of the brain containing cranium).

Actually humans and chimpanzees are descended from a common ancestor - things have moved on since 1927, but the idea is correct, if not in true evolutionary detail.

Skeletons of chimpanzee orang-utan gorilla gcse biology igcse O level revision

Skeletons of a human, chimpanzee, orang-utan and male gorilla in the Natural History Museum, Dublin

 

We (humans and like species, genus Homo) split from chimpanzees (genus Pan) from a common ancestor about 4-7 million years ago - its not that long ago in terms of geological time!

(The Earth is 4500 million years old, 4.5 x 109 years)

The split is difficult to pin-point in time because only fragments of fossil bones are found and they, and the surrounding rock layers, are difficult to date precisely.

Rocks are dated from the ratios of elements in them including lead Pb, samarium Sm, neodymium Nd and argon Ar - the calculations are very complex and not without error, but improving with more research and cross-checking of data between different research groups. This branch of science is called radiometric dating.

 

Human beings and their ancestors are called hominids, but despite the scarcity of good quality fossils, from the fossil record, it is possible to show how we have evolved over the past ~5 million years.

 

The human fossil record and timeline - tabular style for a clear comparison

The early hominid fossils have characteristics somewhere between true apes and true modern humans.

The structure of feet, legs, arms and skull are of particular importance, and of course, a measurement of brain size.

You can also see evidence of evolution through the development of tools using stone and wood.

Gradually, stone tools became more sophisticated, along with brain development.

Generally speaking, the lower the layer of rock the fossils and tools are found in, the older the materials at that level.

The structural features of the tools tells you about their age - the more sophisticated, the more recent.

If a stone tool is found with a wooden handle, or non-mineralised skeleton (still bone), they will contain the element carbon and can be dated by carbon-14 dating (14C is a radioactive isotope of carbon with a half-life of ~5700 years).

For more on dating organic materials see Uses of decay data and half-life values - archaeological radiocarbon dating, dating ancient rocks  GCSE chemistry/physics revision notes

Metals were first extracted and used around 11000 years ago, well after the first appearance of modern humans - copper was extracted from crushed rock and beaten into shape - but smelting came a bit later!

Fossil name and species ~Age in millions of years Skull and brain size (cranial capacity - brain area in skull) Structure of feet Arms and legs Comments
'Ardi' Ardipithecus ramidus female from Ethiopia 4.4 Brain size similar to chimpanzee ~35 kg, 350 cm3 Suggests she climbed trees - had a big toe to grasp branches. She had long arms and short legs - more like an ape than human, but leg structure suggests she walked upright (bipedal) and didn't use hands to walk like apes do. A mixture of ape and human features.
'Lucy' Australopithecus afarensis from Ethiopia 3.2 Brain slightly bigger than Ardi's ~37kg, 430 cm3 but still similar to chimps. Arched feet, more adapted to walking than climbing and no ape-like big toe. The arm and leg sizes are between that of apes and humans, and suggest she walked upright more efficiently than Ardi A mixture of ape and human features, but a bit more 'human' than Ardi.
'Turkana Boy' species of Homo erectus 1.6 Brain much larger than Lucy's,~1000 cm3,  but still less than modern humans Feet and leg structure suggest that he was even better adapted to walking upright than Lucy. He had short arms and long legs, much more human than an ape - better to upright walking than 'Lucy'. Still a mixture of ape and human features, but significantly more 'human-like' than Lucy.
Modern humans Homo sapiens 0.1 - 0.2 significant increase in brain size ~57 kg, 1450 cm3      

 

Further comments on the table of hominids

The table only quotes a few of the many hominid fossils that have been found, but they are, I hope, sufficient, to illustrate the evolutionary path by which a species emerges that can investigate its own complex path of development!

From the table the general evolutionary trends from early hominids to modern humans are:

(i) A general increase in brain size - survival of the best 'thinkers'.

(ii) Arms are no longer required to help in walking - free to develop to do other skilful things!

(iii) Feet and legs evolve to give true bipedal upright walking.

(iv) The craftsmanship and sophistication of stone tool gradually improves.

In the 1980s a scientist called Richard Leakey led an expedition to Kenya in Africa to look for hominid fossils. He and his teams, discovered many important fossils of different species of the genus Australopithecus and genus Homo

Apparently for most of the past 2 million years, there was only a small increase in brain size, but from 100000 to 200000 years ago, evolutionary selection pressures produced a smaller body mass and larger brain.

 

Examples of stone tool development

Homohabilis (2.5 to 1.5 million years ago)

Made simple crude stone tools by bashing rocks together to make sharp flakes that could be used to scrape meat off bone and extract marrow from cracked bones.

Homo erectus (includes Turkana Boy', 2.0 to 0.3 million years ago)

More sophisticated 'sculpturing' of rocks to make simple hand axes - these could be used for chopping meat/wood, scraping meat from bone and a hunting weapon.

Homo Neanderthals (300 000 to 2 million years ago)

A species that went extinct around 30,000 to 40,000 years ago, emerged ~0.4 million years ago.

Able to make good quality flint tools e.g. sharper axes, pointed tools that could bore into wood and wooden spears for hunting.

Flint is a hard sedimentary rock material that can be split into thin layers that can act as blades.

Larger flint tools can be made by striking a large lump to break of flakes and gradually work it into a useful shape like an axe blade - which can be mounted in a wooden shaft.

Homo sapiens (200,000 years ago to the present)

Further improvements in working stone - precisely carved arrow heads from flint.

From 50,000 years ago (at least) a wider range of natural materials is being exploited e.g. spear heads fish hooks from carving antlers, buttons and needles appear in the archaeological record.

 

Some examples of stone tools and dating them

For dating organic materials see archaeological radiocarbon dating, dating ancient rocks  GCSE chemistry/physics revision notes

Radiocarbon dating is based on carbon atoms that were once part of a living organism and so cannot be used to date rock layers or stone tools.

There are several methods used to date rocks in which stone tools are found.

(i) Tools can be put into a crude timeline based on their sophistication.

e.g. as the tools become more recent, their structural features become elaborate.

(ii) Stratigraphy is the study of rock layers.

The deeper the layer of rock where the hominid fossil or stone tool is found in, the older it is more likely to be.

Over time, over thousands of years, any surface layer becomes gradually covered with deposits of e.g. weathered rock or volcanic ash.

This covers fossils and stone tools, entombing them in a particular and successively older geological layer

Therefore you can produce a timeline of hominid fossils and stone tools from what archaeologists and palaeontologists find in success rock layers by (i) digging down carefully through rock layers relatively near the surface or (ii) deeper layers exposed by erosion or tectonic plate activity.

(A palaeontologist is a scientist who studies the fossilized remains of all kinds of organisms (plants, animals, fungi, bacteria and other single-celled living things), and is interested in knowing the history of organic life on earth.)

Derby Museum - Stone tools, mainly flint, used by people 300,000 to 180,000 years ago.

Lincoln

Flint hand axe heads of palaeolithic man from 180,000 t0 450,000 years ago, mostly from Lincolnshire (The Collection Museum, Lincoln)

Some are a bit more sophisticated than the ones in Derby Museum.

Polished stone from the Neolithic Age found on Shetland, much more recent, 4500-6000 years old, much more recent, often produced with great skill, perhaps as ceremonial objects of status?

An elk antler fashioned into a digging tool

An inscribed pendant, 11,000 years old

An efficient barbed hunting too made from antler bone

Flint nodule from smaller flake scraper tools can be made

Examples of Mesolithic ('middle stone age) artefacts in the Yorkshire Museum, York, some from the famous Mesolithic site of Starr Carr near Scarborough. The Mesolithic period in Britain dates from around 11000 to 6000 years ago.

You can see from the series of photographs of stone and wooden tools that they have become more elaborate with decreasing age.

National Archaeology Museum, Dublin, Ireland - well worth a visit - a wonderful museum

Clonycavan Man, an iron age bogman dated, by radiocarbon dating to 392-201 BCE (~25 years old)

 

Changes in the DNA of prehistoric man can be measured with modern genetic analysis, particularly from well preserved bodies found in bogs, usually from some ceremonial-religious ritual activity.

Lactose tolerance and intolerance

For example a mutation occurred, maybe 9000 years ago (the lactose mutation), that allowed neolithic and their modern successors to digest animal milk (cows, goats etc.) beyond breast feeding.

Genetic researchers compared archaeological evidence for 9,000 years of European milk and found an unusually rapid, evolution of lactose tolerance among Europeans well after they first started consuming the animal milk other than human breast milk.

However, some people do not have that particular 'lactose tolerant' mutation and suffer from lactose intolerance, which is quite a common digestive problem where the body is unable to digest lactose, a type of sugar mainly found in milk and dairy products. Symptoms of lactose intolerance usually develop within a few hours of consuming food or drink that contains lactose.


Key points Based on the syllabus-specifications for students taking the AQA, Edexcel and OCR GCSE level biology examinations (~US grades 9-10).

Key points of human evolution

Evidence of Human Evolution

Human evolution is supported by multiple sources of evidence, including fossil skeletons, stone tools, genetic analysis, and other findings.

These clues allow scientists to trace our ancestry, understand adaptations, and determine how humans evolved over millions of years.


1. Fossil Skeletons as Evidence of Human Evolution

Fossils provide physical proof of anatomical changes in hominins over time.

Key Fossil Discoveries

  1. Ardipithecus ramidus (4.4 million years ago)

    • Early hominin with a mix of ape-like and human-like features.

    • Evidence suggests partial bipedalism but still adept at climbing trees.

  2. Australopithecus afarensis ("Lucy") (3.2 million years ago)

    • Walked upright (bipedal) but had a small brain size (~400cc).

    • Foot structure suggests human-like movement, proving early adaptation to walking.

  3. Homo habilis (2.4–1.4 million years ago)

    • Known as "Handy Man" because of simple stone tool use.

    • Larger brain (~600cc) compared to earlier hominins, indicating cognitive development.

  4. Homo erectus (1.8 million – 300,000 years ago)

    • Fully upright posture and larger brain (~1000cc).

    • Used advanced stone tools and fire, showing problem-solving skills.

  5. Homo neanderthalensis (400,000 – 40,000 years ago)

    • Stocky build adapted for cold climates.

    • Evidence of social structures, tool-making, and burial practices.

  6. Homo sapiens (300,000 years ago – present)

    • Larger brain (~1300cc) and complex reasoning skills.

    • Development of language, culture, and advanced technology.


2. Stone Tools as Evidence of Cognitive Evolution

Stone tools illustrate the increasing intelligence and problem-solving abilities of human ancestors.

Progression of Tool Use

  • Oldowan Tools (Homo habilis, ~2.5 million years ago)

    • Simple stone flakes used for cutting meat.

  • Acheulean Tools (Homo erectus, ~1.6 million years ago)

    • Hand axes for hunting and preparing food.

  • Mousterian Tools (Neanderthals, ~200,000 years ago)

    • More advanced and specialized tools, suggesting planning.

  • Upper Paleolithic Tools (Homo sapiens, ~40,000 years ago)

    • Needles, fishhooks, and finely carved weapons for complex survival strategies.


3. Genetic Analysis as Evidence of Evolution

Modern genetic research helps confirm relationships between ancient hominins and present-day humans.

Key Genetic Evidence

  • Comparative DNA studies show humans share 98–99% of DNA with chimpanzees, confirming common ancestry.

  • Neanderthal DNA: Some modern humans (European and Asian populations) contain ~1-2% Neanderthal genes, showing interbreeding occurred.

  • Mitochondrial DNA (mtDNA) supports the "Out of Africa" theory, tracing all modern human ancestry to a single African population.


4. Other Evidence Supporting Human Evolution

  • Fossilized Footprints: Evidence from Laetoli, Tanzania (~3.6 million years ago) shows early bipedalism in Australopithecus.

  • Brain Size Increase: Skull fossils show how brain size expanded over time, allowing advanced thought.

  • Cultural Evolution: Cave paintings, symbolic objects, and burial rituals highlight growing intelligence and abstract thinking.


Importance of These Findings on human evolution

  1. Confirms Common Ancestry – Fossil and genetic evidence proves humans evolved from ape-like ancestors.

  2. Explains Adaptations – Shows how walking, tool use, and brain development allowed survival.

  3. Supports Evolutionary Theory – Provides clear proof that natural selection shaped human traits over time.

The combined study of fossils, tools, genetics, and cultural artifacts creates a comprehensive picture of human evolution, helping us understand our origins and how we became the species we are today.


Summary of learning objectives and key words or phrases

Know and be able to describe or interpret, evidence of human evolution from the fossil record of skeletons.

Know how stone tools, bone tools and modern genetic analysis all contribute to understand the evolution of human beings.

Know about the lactose tolerance mutation that occurred in prehistoric man.

Be able to discuss and interpret archaeological findings of palaeolithic man, neolithic man and bronze age man.


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